Board Thread:Lore Discussion/@comment-2165692-20161203091356/@comment-26893431-20161203130139

I think the thumb would be like the Night Mother and her nail would be the Listener, while the palm of the hand would represent Sithis, since he is connected to all.

That's just my theory though.

As for the Keeper, it might be (contradicting my own theory already, great!) that the Listener is the Thumb and the Keeper is his Nail. And that Sithis and the Night mother are the Palm of the Hand.

Again, just a theory.